FABACEAE (LEGUMINOSAE) [Draft]

Key to subfamilies

1a.       Flowers actinomorphic, petals valvate in bud, free or synpetalous; anthers sometimes with a deciduous gland at apex ................................................................................................  1. Mimosoideae

1b.       Flowers more or less zygomorphic, petals imbricate in bud.

2a.       Flowers slightly zygomorphic; corolla not papilionaceous, adaxial (uppermost) petal overlapped on each side by the adjacent lateral petals (when these present); stamens with usually free filaments .  2. Caesalpinioideae

2b.       Flowers strongly zygomorphic (very rarely actinomorphic); corolla papilionaceous, adaxial petal (vexillum) outside the adjacent lateral petals (wing), abaxial petals (keel) basally connate, stamens diadelphous (9+1) or monadelphous, rarely free ....................................................................  3. Papilionoideae

1. Subfam. MIMOSOIDEAE

Key to genera

1a.       Stamens 10 or less.

2a.       Climbers or herbs.

3a.       Woody climbers; leaves with pinnae 2 pairs, tenderils present; pods jointed ........  3. Entada

3b.       Perennial herbs; leaves with pinnae 4–10 pairs, without tendrils; pods not jointed  4. Neptunia

2b.       Trees, shrubs or subshrubs.

4a.       Tall trees, 6–30 m tall.

5a.       Pinnae with leaflets 28–60 pairs; flowers in club-shaped pendulous head; pods base attenuate into a long stipe .........................................................................................................  1. Parkia

5b.       Pinnae with leaflets 4–7 pairs; flowers in racemes or panicles; pods base not attenuate into a long stipe ..............................................................................................................  2. Adenanthera

4b.       Small trees, shrubs or subshrubs, 0.5–6 m tall.

6a.       Armed plants; pods dehiscent in several segmens separating from persistent sutures; leaflets sensitive ......................................................................................................................  6. Mimosa

6b.       Unarmed plants; pods not dehiscent in several segments separating from persistent sutures; leaflets not sensitive.

7a.       Small trees or shrubs, 2–6 m tall; pinnae with leaflets 5–15 pairs; pods broadly linear  5. Leucaena

7b.       Subshrubs, 0.5–1.3 m tall; pinnae with leaflets 6–21 pairs; pods linear
                                                                                                               7. Desmanthus

1b.       Stamens numerous, usually more than 10.

8a.       Filaments free or only connate at base ...................................................................  8. Acacia

8b.       Filaments connate into a tube.

9a.       Pods dehiscent into 2 valves.

10a.     Pods curved into a circle; stipules spinnescent ......................................  9. Pithecellobium

10b.     Pods straight or slightly curved; stipules not spinnescent.

11a.     Pods with thickened sutures, elastically dehiscing from apex and downwards; seeds with pleurogram                                                                                                                 10. Calliandra

11b.     Pods without thickend sutures, dehiscing along both dorsal and ventral sutures; seeds without pleurogram ........................................................................................  11. Archdendron

9b.       Pods indehiscent or tardy dehiscent.

12a.     Pods contorted into a half or full circle .................................................  12. Enterolobium

12b.     Pods straight.

13a.     Flowers in spikes ...................................................................................  13. Falcataria

13b.     Flowers in head or short corymbs.

14a.     Pods septate between seeds, thick, fleshy ..............................................  14. Samanea

14b.     Pods not septate between seeds, flat, thin ..................................................  15. Albizia

2. Subfam. CAESALPINIOIDEAE

Key to genera

1a.       Leaves simple, entire or bifid to bi-lobed, palmately nerved.

2a.       Stamens 10; flowers purple-red or pink; pods narrowly winged on adaxial suture .  14. Cercis

2b.       Stamens 3 or 5, if 10, then flowers white or yellowish or green; pods not winged  15. Bauhinia

1b.       Leaves once pinnate or bipinnate.

3a.       Leaves once pinnate.

4a.       Petals absent.

5a.       Flowers smaller, red; leaflets 2–4 pairs ......................................................  11. Ceratonia

5b.       Flowers larger, yellow or dark red; leaflets 4–6 pairs ......................................  17. Saraca

4b.       Petals present.

6a.       Petals 1–3.

7a.       Petals 3; seeds without aril ...................................................................  21. Tamarindus

7b.       Petals 1(2); seeds arillate.

8a.       Petals long clawed; stamens 7–8 ...............................................................  18. Afzelia

8b.       Petals sessile; stamens 9 ..........................................................................  20. Sindora

6b.       Petals 5.

9a.       Leaves imparipinnate; leaflets alternate; pods narrowly winged on sutures
                                                                                                                        12. Zenia

9b.       Leaves paripinnate; leaflets opposite; pods not winged.

10a.     Leaflets 1-paired .................................................................................  19. Hymenaea

10b.     Leaflets 3–5-paired.

11a.     Flowers yellow; stamens 4–10 ................................................................  13. Cassia

11b.     Flowers purple or pink; stamens 2 .........................................................  16. Lysidice

3b.       Leaves bipinnate

12a.     Flowers polygamous; pods mostly pulpy within between seeds.

13a.     Branches not spinose; leaflets entire; pods turgid or subterete, 2-valved
.............................................................................................................  1. Gymnocladus

13b.     Branches and trunk armed with simple or branched spines; leaflets often ± serrate; pods compressed, indehiscent or tardily 2-valved ......................................................................  2. Gledetsia

12b.     Flowers bisexual.

14a.     Tall trees, unarmed.

15a.     Flowers large, very showy, more than 7 cm in diam.; pods loriform, large, subligneous  5. Delonix

15b.     Flowers small or medium, not more than 3 cm in diam.

16a.     Flowers scarlet; stamens 5, pods long-stipitate, narrowly winged along adaxialy suture  3. Acrocarpus

16b.     Flowers yellow or yellowish-green; stamens 10; pods winged along both sutures or not winged.

17a.     Pods winged along both sutures; flowers over 15 mm in diam.; stamens slightly shorter than petals .........................................................................................................  4. Peltophrum

17b.     Pods not winged along sutures; flowers below 5 mm in diam.; stamens double longer than petals ...................................................................................................  10. Erythophleum

14b.     Climbers or small trees, shrubs, armed.

18a.     Flowers subregular, nearly actinomorphic.

19a.     Climbers; pods samaroid, indehiscent; leaflets 6–30-paired .................  7. Pterolobium

19b.     Trees; Pod not samaroid, opening along middle of valves; leaflets 2–4-paired  8. Haematoxylon

18b.     Flowers irregular, zygomorphic.

20a.     Leaf-rhachis ending in a spine-like sharp point, leafless very small; pods torulose  9. Parkinsonia

20b.     Leaf-rhachis not ending in a spine-like sharp point; leaflets not very small; pods not torulose, ovate to oblong, or falcate ............................................................................  6. Caesalpinia

3. Subfam. FABOIDEAE (PAPILIONOIDEAE)

Key to tribes

1a.       Filaments all free or connate only at base, anthers uniform.

2a.       Leaves odd-pinnate (simple leaf only in Bowringia calicarpa and Ormosia simplicifolia); stipules small or absent; calyx usually subequally 5-dentate; trees or shrubs, rarely vines ........  1. Sophoreae

2b.       Leaves palmate-trifoliolate (rarely 1-foliolate in Ammopiptanthus mongolicus); stipules usually united with petioles or amplexical, without stipels; calyx usually deeply 5-lobed; shrubs or herbs .  21. Thermopsideae

1b.       Filaments partly or almost wholly united to one another, either monadelphous in a closed tube, or diadelphous, in the latter case the vexillary (adaxial) filament often free or partly free from the remainder; anthers uniform or dimorphic.

3a.       Anthers uniform in size and shape or nearly so, not alternately basifixed and dorsifixed, also not alternately longer and shorter.

4a.       Free upper part of filaments more or less filiform, not dilated upwards.

5a.       Pods not breaking up into separate segments when mature; seeds 1 to several.

6a.       Stamens monadelphous and filaments partly connate into a single more or less closed tube, sometimes the adaxial filament free at the base but connate higher up with the remainder.

7a.       Herbs with indumentum composed mainly of medifixed (T-shaped) hairs, racemes axillary, anther-connective apiculate; style glabrous
.....................................................................................  6. Indigofereae (Cyamopsis)

7b.       Trees or shrubs, rarely herbs, indumentum when present not composed of medifixed (T-shaped) hairs; anther-connective not apiculate; inflorescens racemose or pinnulate, or flower solitary; style glabrous or hairy.

8a.       Pods indehiscent; semi-orbicular or circinate, 1–2-seeded; annual or perennial herbs  13. Hedysareae (Onobrychis)

8b.       Pods dehiscent, not semi-orbicular.

9a.       Trees, shrubs , sometimes climbing; stipules not sagittate or absent
                                                                                                 4. Millettieae (p.p.)

9b.       Herbs; stipules sagittate .....................................................  12. Galegeae (Galega)

6b.       Stamens diadelphous, mostly vexillary filament free or partly adnate to the remainder, or if monadelphous then free at upper part or top.

10a.     Leaves imparinnate or pinnately or digitately 3-foliolate, sometimes 1-foliolate (Tadehagi) or primitively simple (Indigofera spp.).

11a.     Leaves gland-dotted or pellucide-punctate below.

12a.     Pods dehiscent, 1-several-seeded; leaves trifoliolate or simple (Flemingia spp.)  8. Phaseoleae (Cajaninae)

12b.     Pods indehiscent, 1-seeded; leaves simple ........................................  9. Psoraleeae

11b.     Leaves neither gland-dotted n or pellucide-punctate below.

13a.     Plant with indumentum composed mainly of T-shaped hairs; racemes or spikes axillary; anther-connective tipped by a gland or mucro
                                                                                                        6. Indigofereae

13b.     Plant glabrous or without indumentum composed of T-shaped hairs; anthers neither tipped by a gland nor mucronate.

14a.     Leaflets more or less toothed, lateral nerves continued to the margin; mostly herbs;  stipules often adnate to the petiole; stipe absent ..........................................  18. Trifolieae

14b.     Leaflets entire, lateral nerves not continued to margin; mostly shrubs or subshrubs, rarely herbs; stipellate.

15a.     Pods jointed but dehiscing its full length .............................  7. Desmodieae (p.p)

15b.     Pods not jointed ................................................................  8. Phaseoleae (p.p.)

10b.     Leaves pinnate (including pinnately 3-foliolate).

16a.     Leaves gland-dotted or pellucide-punctate below; ovules few.

17a.     Pods dehiscent; leaves pinnate-trifoliolate; corolla regular ........  8. Phaseoleae (p.p.)

17b.     Pods indehiscent;leaves imparipinnate; corolla without wings and keel
                                                                                                       10. Amorpheae

16b.     Leaves not gland-dotted or pellucide-punctate below; ovules usually more than 4, rarely 2 or 1.

18a.     Pods indehiscent; without stipels.

19a.     Pods inflated and blader-like .......................................................  12. Phaseoleae

19b.     Pods not inflated and blader-like

20a.     Leaves imparipinnate; leaflets alternate .......................................  2. Dalbergieae

20b.     Leaves trifoliolate; 1-seeded.

21a.     Pods plano-compressed, leathery .................  7. Desmodieae (Lespedezineae)

21b.     Pods drupe-like, fragile .........................................................  20. Euchresteae

18b.     Pods dehiscent, sometimes only at apex.

22a.     Petals very unequal in length, either vexillum or the keel petals much longer than other; stipels present, sometimes gland-like
.............................................................................  3. Phaseoleae (Erythrininae)

22b.     Petals more or less equal in length (or at any rate the keel-petals not much longer than others, though sometimes beaked or spirally twisted).

23a.     Indumentum (when present) usually composed, at least in part, of adpressed medifixed (T-shaped) hairs; anther-connective with a gland or mucro at apex; stipels present or not obvious
                                                                               6. Indigofereae (Indigofera)

23b.     Indumentum not as above; anther-connective without a gland or mucro at the apex.

24a.     Rhachis of inflorescence thickened (nodose) at insertion of pedicels, mostly stipellate       8. Phaseoleae

24b.     Rhachis of inflorescence not thickend at insertion of pedicels; leaves without stepel.

25a.     Rhachis of leaves transformed at apex into a tendril or bristle; stipels absent.

26a.     Stamens 10; flowers solitary, fasciculate or racemose in leaf-axils; claw of vexillum free from staminal-sheath ................................................................  16. Fabeae

26b.     Stamens 9, flowers in terminal or subterminal racemes; claw of vexillum more or less adnate to staminal-sheath ...........................................................  3. Abreae

25b.     Rhachis of leaves not ending in a tendril or bristle.

27a.     Pods longitudinally septate within by the intrusion of seed-bearing suture; style glabrous .....................................................................  12. Galegeae (Astragalinae)

27b.     Pods not longitudinally separate as described above, sometimes transversely septate but not breaking up into separate joints.

28a.     Pods inflated and bladder-like, sometimes dehiscent only at the apex  12. Galegeae (p.p.)

28b.     Pods compressed.

29a.     Stipules mostly adnate to the petiole; leaflets often toothed; mostly herbs  18. Trifolieae (p.p.)

29b.     Stipules not adnate to the petiole, or absent; leaflets entire; woody or herbs.

30a.     Ovary 1-seeded ..........................................................  7. Desmodieae

30b.     Ovary 2 to many seeded.

31a.     Flowers in terminal or leaf-opposed racemes, rarely axillary  4. Millettieae (p.p.)

31b.     Flowers single, fasciculate or in racemes axillary.

32a.     Erect plants; leaves paripinnate or imparipinnate; leaflets 2–4 pairs  5. Robinieae (Sesbania)

32b.     Twining herbs; leavespinnately or digitately 3-foliolate
......................................................................  8. Phaseoleae (p.p.)

5b.       Pods breaking up into 1-seeded segments when ripe.

33a.     Leaves with stupels .......................................................  7. Desmodieae (Desmodiinae)

33b.     Leaves without stipels.

34a.     Keel-petals often obliquely truncate at the apex; wings short or very small, rarely equalling the keel-petals ......................................................................................  13. Hedysareae (p.p.)

34b.     Keel-petals obtuse or breaked, incurved; wings often transversely plicate
.................................................................................................  11. Aeschynomeneae

4b.       Free upper part of all or half of the filaments dilated or expanded upwards.

35a.     Pods jointed and breaking up into separate segments, more or less tetragonous, indehiscent       15. Coronilleae

35b.     Pods 2-valved or indehiscent, sometimes transversely septate but not breaking up into separate joints.

36a.     Leaf-rachis ending in a bristle or tendril; flowers solitary or racemose in the leaf-axil  17. Cicereae

36b.     Leaf-rachis not ending in a bristle or tendril.

37a.     Claws of some or all of the petals partly adnate to the staminal sheath; leaflets usually toothed, the lateral nerves extended to the teeth or margin
...................................................................................................  18. Trifolieae (p.p.)

37b.     Claws of the petals not adnate to the staminal sheath.

38a.     Stipules gland-like, free from the petiuole; leaflets entire, nerves not running into the margin; lowermost pair often resembling stipules ..................................................  14. Loteae

38b.     Stipules mostly partly adnate to the petiole; nerves of the leaves extended to the usually toothed margin ......................................................................................  18. Trifolieae (p.p.)

3b.       Anther dimorphic, alternately dorsifixed and basifixed, either all equal or alternately longer and shorter.

39a.     Upper part of filaments swollen or expanded; perennial of shrublets, often spiny and glandular       18. Trifolieae (Ononis)

39b.     Upper part of filaments neither swollen nor expanded.

40a.     Pods transversely septate and breaking up into 1-seeded joints (sometimes some fruits reduced to 1 joint) ...................................................................................  11. Aeschynomeneae (p.p.)

40b.     Pods not transversely septate or if so then not breaking up into separate joints.

41a.     Filaments connate into a more or less closed tube, but free at the top; stipel absent or very small.

42a.     Claws of the lower petals more or less adnate to the staminal tube; stipules minute or absent        20. Genisteae

42b.     Claws of all the parts free from the staminal tube.

43a.     Ovary 1-ovuled; pods 1-seeded ...................................  11. Aeschynomeneae (p.p.)

43b.     Ovary more than 1-ovuled; pods more than 1-seeded ...............  20. Genisteae (p.p.)

41b.     Filaments connate into a sheath split adaxially or sometimes the adaxially (vexillary) filament free or partly so.

44a.     Leaves simple; flowers mostly in terminal or leaf-opposed racemes or spikes; style bearded        19. Crotalarieae (p.p.)

44b.     Leaves 3-or more-foliolate.

45a.     Leaves glandular below; style bearded lengthwise or pilose around the stigma  8. Phaseoleae (p.p.)

45b.     Leaves not glandular below; style glabrous or hairy.

46a.     Style glabrous.

47a.     Leaves pinnately foliolate; stipels mostly present; mostly climbers; pods sometimes ribbed, often clothed with stinging hairs; inflorescence axillary or born on old stem  8. Phaseoleae (Mucuna)

47b.     Leaves digitately 3-foliolate; stipels absent; creeping herbs; pods without shining hairs; inflorescence mostly terminal or leaf-opposed
.............................................................................  19. Crotalarieae (Lotononis)

46b.     Style hairy.

48a.     Style bearded lengthwise; pods turgid or inflated ..............  19. Crotalarieae (p.p.)

48b.     Style at most pubescent only towards the base; pods not inflated
................................................................................  8. Phaseoleae (Glycininae)

1. Tribe Sophoreae

1a.       Pods terete, moniliform, rarely 4-winged (S. mollis) ..............................................  4. Sophora

1b.       Pods compressed or turgid, sometimes winged along sutures.

2a.       Leaves simple, scandent shrubs; corolla white; pods ovoid, transversely nerved
.........................................................................................................................  6. Bowingia

2b.       Leaves pinnate (except O. simplicifolia), erect shrubs or small trees; corolla pink; pods long elliptic or obovate, not transversely nerved.

3a.       Leaf-rachis ending in a spine; indument silvery-silky ................................  5. Ammodendron

3b.       Leaf-rachis not ending in a spine; indument not silvery-silky.

4a.       Pods compressed or more or less turgid; sutures neither winged nor thickened  1. Ormosia

4b.       Pods complanate, sutures slightly thickened on one side or both side.

5a.       Buds of shoots embraced by the inflated base of the petiole ......................  2. Cladrastis

5b.       Buds of shoots not embraced by the inflated base of the petiole ...................  3. Macckia

2. Tribe Dalgergieae

1a.       Flowers white or red; pods oblong or linear.......................................................... 7. Dalbergia

1b.       Flowers yellow; pods orbicular......................................................................... 8. Pterocarpus

3. Tribe Abreae

Only one genus Abrus.

4. Tribe MILLETTIEAE

1a.       Herbs or shrublet-like; stipules present .............................................................  19. Tephrosia

1b.       Woody plants; stipules absent or present.

2a.       Stipules absent.

3a.       Trees; pods thickly woody, convex, 1-seeded ......................................  10. Antheroporum

3b.       Vines or climber; pods compressed.

4a.       Pods thick, not winged .............................................................................  14. Pongamia

4b.       Pods thin, winged ...........................................................................................  18. Derris

2b.       Stipules present, deciduous or persistent.

5a.       Deciduous vines with terminal pendulous racemes ...........................................  15. Wisteria

5b.       Evergreen vines with terminal or axillary panicles or racemes, sometimes cauliflorous.

6a.       Inflorescence axillary or cauliflorous; ovary 2-ovuled.

7a.       Stipels present; flowers small, fasciculate; pods leathery, compressed ...........  12. Fordia

7b.       Stipels absent; flowers large, solitary; pods woody, turgid (Whitfordiodendron in FRPS)        11. Afgekia

6b.       Inflorescence terminal or axillary; ovary 4 or more-ovuled.

8a.       Pods narrowly winged; leaves 3-foliolate ......................................  17. Craspedolobium

8b.       Pods not winged; leaves imparipinnate, more than 5-foliolate.

9a.       Exocarp separated from endocarp when dry ........................................  16. Sarcodum

9b.       Exocarp not separated from endocarp when dry (Note! Millettia sect. Eurybotryae Dunn = Callerya Endlicher (see Legumes World, 370. 2005). What opinion have the authors of FOC?) ...............................................................................................................  13. Millettia

5. Tribe Robinieae

1a.       Flowers fasciculate at ends of shoots; leaflets linear and soon folded; grown in highland  20. Salweenia

1b.       Flowers in racemes; leaflets oblong to broad-ovate; grown in lowland.

2a.       Trees or shrubs; leaflets 2–12 pairs; stipules spinescent; pods flat and compressed, narrowly winged along the upper suture, continuous inside ............................................................  21. Robinia

2b.       Herbs or shrub-like herbs; leaflets 10–32 pairs; stipules not spinescent; pods, linear and subterete not winged along the suture, septate between seeds inside
........................................................................................................................  22. Sesbania

6. Tribe Indigofereae

1a.       Stamens diadelphous; vexillum hairy outside, if not hairy than pods 1-angulate
                                                                                                                                                       23. Indigofera

1b.       Stamens uniform; vexillum not hairy outside; pods subtetragonous-compressed
                                                                                                                                                       24. Cyamopsis

7. Tribe Desmodieae

1a.       Stipels absent; pods 1-jointed, 1-seeded, not glochidiate.

2a.       Twining herbs; bracts membranous, enlarged after flowering, leaf-like, folded
.....................................................................................................................  39. Phylacium

2b.       Shrubs or herbs, not twining, bracts not like above..

3a.       Lateral veins of leaflets strict; stipules large .............................................  42. Kummerowia

3b.       Lateral veins of leaflets arch-curved near margin; stipules small, subulate.

4a.       Bracts caduceus, 1-flowered; pedicels articulate below calyx; keel falcate, acute  40. Campylotropis

4b.       Bracts persistent, 2-flowered; pedicels not articulate; keel strict, obtuse
.................................................................................................................  41. Lespedeza

1b.       Stipels present; pods usually glochidiate, 2–several-jointed, rarely 1-jointed, 1-seeded.

5a.       Branch nodes with trifid and hard spines; leaves 1-foliolate ......................  25. Trifidacanthus

5b.       Branch nodes without trifid and hard spines; leaves 3(–9)-foliolate, rarely 1-foliolate.

6a.       Calyx glume-like, lobes dry and hard, striate ..............................................  38. Alysicarpus

6b.       Calyx not glume-like, lobes herbaceous, not striate.

7a.       Umbels or short racemes axillary; flowers white or yellow; leaves 3-foliolate.

8a.       Umbels not enclosed in 2 opposite leafy bracts .................................  26. Dendrolobium

8b.       Umbels enclosed in 2 opposite leafy bracts ...........................................  27. Phyllodium

7b.       Racemes or panicles terminal or axillary, rarely flowers fasciculate axillary; leaves (1)3–9-foliolate.

9a.       Leaves digitately 3-foliolate;stipules united, apex 3-lobed; pods 2-jointed
                                                                                                                   28. Dicerma

9b.       Leaves pinnately (1)3–9-foliolate; stipules free, apex entire; pods 2–several-jointed, rarely 1-jointed.

10a.     Pods distinctly stipitate, ventral suture deeply incrised, the isthmus less than 1/5 as broad as pod; stamens monadelphous (Podocarpum in FRPS)
........................................................................................................  30. Hylodesmum

10b.     Pods not stipitate or rarely shortly stipitate; constricted on both sutures or ventral suture straight; stamens diadelphous, rarely monadelphous.

11a.     Joints of pod plicate-retrofracted.

12a.     Calyx lobes short and broad, enlarged after flowering, net-veined .........  37. Christa

12b.     Calyx lobes subulate or setaceous, not englarged after flowering, net-veined.

13a.     Joints connected along ventrual suture .................................................  35. Uraria

13b.     Joints connected at center of the joint (Note! should Urariopsis be combined with Uraria?) ....................................................................................................  36. Urariopsis

11b.     Joints of pod not plicate-retrofracted.

14a.     Pedicels hooked-inflexed at apex and inverting flowers; leaves 1-foliolate  34. Mecopus

14b.     Pedicel not hooked-inflexed at apex.

15a.     Petioles winged; leaves 1-foliolate ..................................................  33. Tadehagi

15b.     Petioles not winged, if winged, then 3-foliolate.

16a.     Pods turgid, continuous within, not jointed, valves transversely lined-venose  32. Pycnospora

16b.     Pods not turgid, jointed, valves not transversely line-venose.

17a.     Joints of pod indehiscent ........................................................  29. Desmodium

17b.     Joints of pod dehiscing along dorsal suture, remaining attached to other suture  31. Codariocalyx

8. Tribe Phaseoleae

1a.       Leaflets and calyx generally with yellowish gland-dots; bracteoles absent; inflorescences not nodose (7. subtribe Cajaninae).

2a.       Pods 3- or more-seeded.

3a.       Pods depressed between seeds ......................................................................  74. Cajanus

3b.       Pods not depressed between seeds ..............................................................  75. Dunbaria

2b.       Pods 1- or 2-seeded.

4a.       Leaves 1-foliolate; funicle attached at end of linear hilum ...............................  78. Eriosema

4b.       Leaves digitately or pinnately 3-foliolate, or simple (not 1-foliolate); funicle attached at middle of hilum.

5a.       Erect herbs or shrubs; leaves digitately 3-foliolate or simple; pods turgid
.................................................................................................................  76. Flemingia

5b.       Twining herbs or shrublets; leaves pinnately 3-foliolate; pods compressed
...............................................................................................................  77. Rhynchosia

1b.       Leaflets and calyx without gland-dots.

6a.       Style flattented, thicked or twisted, mostly barbate, if not barbate and terete, then vexillum and keels with minute appendages; hilum usually covered with spongy tissue (6. subtribe Phaseolinae).

7a.       Style flattened laterally; vexillum with a single large appendage............................ 68. Lablab

7b.       Style terete.

8a.       Keel-petals apex spirally twisted.

9a.       Wing-petals longer than vexillum; style twice 90Ί incurved; pods 2–3mm broad  72. Macroptilium

9b.       Wing-petals shorter than vexillum; style 360Ί or more twisted; pods 1–1.5 cm broad  73. Phaseolus

8b.       Keel-petals apex not spirally twisted.

10a.     Pods 4-winged ................................................................................  66. Psophocarpus

10b.     Pods not 4-winged.

11a.     Stigma lateral .............................................................................................  71. Vigma

11b.     Stigma terminal.

12a.     Vexillum appendages absent; corolla purple-blue .............................  65. Dysolobium

12b.     Vexillum appendages present.

13a.     Vexillum appendages short; corolla white or violet ..............................  69. Dolichos

13b.     Vexillum appendages long and narrow; corolla light yellow-green
                                                                                                    70. Macrotyloma

6b.       Style generally terete and unbearded (sometimes a few hairs belowe stigma), occasionally coiled (in Erythrininae), sometimes bearded or flattened (in Clitoriinae); hilum rarely covered with spongy tissue (some Erythrina except).

14a.     Flowers generally resupinate; calyx naked inside; style narrowed, penicillate or bearded distal part; corolla often hairy; leaflets 3, 1 or 5–9, with minute hooked hairs (5. subtribe Clitoriinae).

15a.     Vexillum with a conspicuous spur on the back ........................................  63. Centrosema

15b.     Vexillum without spur on the back ................................................................  64. Clitoria

14b.     Flowers not resupinate or if so differing in other respects.

16a.     Vexillum silky hairy outside, rather small, without appendages inside; seeds smooth, with a prominent aril; inflorescence not or only slightly nodose (4. subtribe Ophrestiinae) .....  62. Ophrestia

16b.     Vexillum glabrous or if hairy then inflorescence generally nodose or flowers much modified.

17a.     Flowers mostly adapted to birds or bats for pollenation, petals generally unequal in length, sometimes with small bee-type flowers but then either scoiled (Apios, Cochlianthus) or flowers in extensive panicles and pod samaroid (Spatholobus) (1. subtribe Erythrininae).

18a.     Trees or shrubs; stems with prickles; keel-petals much shorter than the vexillum  43. Erythrina

18b.     Climbers, rarely trees (Butea monosperma); keel-petals usually the largest petals; stems without prickles.

19a.     Style spirally coiled.

20a.     Leaves 3-foliolate, turns black on drying .....................................  48. Cochlianthus

20b.     Leaves 3–7-foliolate, remains green on drying .........................................  47. Apios

19b.     Style not spirally coiled.

21a.     Pods many-seeded, 2-valved; stinging hairs present .............................  44. Mucuna

21b.     Pods 1-seeded at the apex, empty below; stinging hairs absent.

22a.     Flowers yellow to scarlet, 1.5–8 cm .....................................................  45. Butea

22b.     Flowers purple, rose, or white, 5–10 m .....................................  46. Spatholobus

17b.     Flowers mostly adapted to bees for pollination or if bird-flowers then petals subequal in length.

23a.     Inflorescences generally nodose, seeds diverse, with short to long hilum (2. subtribe Diocleinae).

24a.     Stigma lateral to subterminal; ovary scarcely pilose, hairs extending up style, forming a falsbeard .......................................................................................................  50. Pachyrhizus

24b.     Stigma terminal; style nacked.

25a.     Calyx 2-lipped, the upper lip large, entire or 2-lobed, the lower lip small; stamens monodelphous; pods loriform or oblong ..........................................  49. Canavalia

25b.     Calyx not strongly 2-lipped; stamens diadelphous; pods linear.

26a.     Upper calyx-lobes completely united (calyx appearing 4-lobed) .......  51. Galactia

26b.     Upper calyx-lobes slightly to largely separate .........................  52. Calopogonium

23b.     Inflorescences not or scarcely nodose; seeds smooth or granular, with short hilum (3. subtribe Glycininae)

27a.     Pods indehiscent; seeds with filiform funicles .....................................  59a. Mastersia

27b.     Pods dehiscent.

28a.     Anthers alternately small and sterile; persistent style of pods turned up into a hook  57. Teramnus

28b.     Anthers all fertile; pods unhooked.

29a.     Stems strongly 4-angled, from the corners of which descend brown hairs; flowers ca. 5 mm ........................................................................................................  58. Teyleria

29b.     Stems not strongly 4-angled, and without brown hairs on the corners; flowers longer than 5 mm.

30a.     Claws of the wings and keel shorter than the blade; seeds usually rough-sufaced; hilum often with a scarious strophiole ovary-walls opaque.

31a.     Leaves 1-foliolate ..........................................................................  54. Nogra

31b.     Leaves 3-foliolate.

32a.     Flowers 2 or more per node of the inflorescence ......................  53. Pueraria

32b.     Flowers 1 per node of the inflorescence.

33a.     Flowers 12 mm or more ..................................................  55. Sinodolichus

33b.     Flowers 9 mm or less ..............................................................  56. Glycine

30b.     Claws of wings and keel longer than the blade; seeds smooth surfaced, without a scarious strphiole; ovary-walls often transparent.

34a.     Calyx-lobes truncate; flowers yellow ..........................................  60. Dumasia

34b.     Calyx-lobes deltoid, evident; flowers usually red-purple, purple, blue or white, never yellow.

35a.     Pods septate between seeds;vexillum suberect ..........................  59. Shuteria

35b.     Pods continuous within; vexillum erect .............................  61. Amphicarpaea

9. Tribe Psoraleeae

Only one genus: Psoralea.

10. Tribe Amorpheae

Only one genus: Amorhpa.

11. Tribe Aeschynomeneae

1a.       Leaves, calyx, etc with pellucid glands, if glands obscure then flowers sessile in numerous paired stipule-like bacts; leaves 2-foliolate .........................................................................................  85. Zornia

1b.       Leaves, calyx, etc. without pellucid glands or sometimes with dark dots; flowers sessile or pedicellate.

2a.       Flowers sessile or subsessile; receptacle longer than broad, sometimes tubular; ovary sessile or subsessile; tannins in mesophyll forming reticulate patterns of dry leaflets abaxially (4. subtribe Stylosanthinae).

3a.       Leaves 3-foliolate; pods 1–2-jointed, not developing underground ...........  86. Stylosanthes

3b.       Leaves 4-foliolate; pods subtorulose but not developing underground ...............  87. Arachis

2b.       Flowers pedicellate; receptacle broader than long or o [=??]; ovary stipitate; tannins in mesophyll not forming reticulate patterns of dry leaflets.

4a.       Leaves imparipinnate; pods with strong continuous nerves on sides; ovary-septa formed late; stamens monadelphous (1. subtribe Ormocarpinae)
                                                                                                                81. Ormocarpum

4b.       Leaves paripinnate,or occasionally imparipinnate; articles of pod reticulately veined; ovary-septa formed by anthesis; stamens monadelphous or diadelphous (5+5) (2. subtribe Aeschynomeninae).

5a.       Bracts much enlarged, usually completely enveloping the flowers and fruits; pods 1–2-jointed     84. Geissaspis

5b.       Bracts not completely enveloping the flowers and fruits; pods 1–more-jointed.

6a.       Inflorescencess mostly lax; pods exserted from the calyx ..................  82. Aeschynomene

6b.       Inflorescences mostly dense and ± scorpioid; pods enclosed by the persistent and accrescent calyx                                                                                                                      83. Smithia

12. Tribe Galegeae

1a.       Style bearded, sometimes just a tuft of hairs below stigma on one side (1. subtribe Coluteinae).

2a.       Leaves reduced to scales; flowers single; pods compressed .....................  90. Eremosparton

2b.       Leaves imparipinnate, 7–25-foliolate; flowers in racemes; pods inflated.

3a.       Flowers yellow; vexillum with 2 callosities above the claw ................................  88. Colutea

3b.       Flowers red to violet; vexillum without callosities above claw .................  89. Sphaerophysa

1b.       Style glabrous, but stigma sometimes penicillate (sometimes the inner side of style upper part hairy in Astragalus).

4a.       Anther-thecae confluent towards apex; leaves pulvinate, spirally arranged; some hairs glandular-peltate or -capitate (4. subtribe Glycyrrhizinae) .......................................................  102. Glycyrrhiza

4b.       Anther-thecae separate; leaf-pulvinus reduced or absent; without peltate glandular hairs.

5a.       Pod-nerves transverse, confluent over middle part of valves, fine or inconspicuous; vexillary filament generally free, keel-petals generally auriculate; leaflet-nerves usually looped within the margin or inconspicuous (2. subtribe Astragalinae).

6a.       Calyx base usually oblique, upper side ± slightly gibbous; wings pinnately nerved; pod valves general twisting on deshiscence.

7a.       Leaves paripinnate, rhachis persistent and hardened into a spines.

8a.       Pods broadly inflated; flowers violet ............................................  91. Halimodendron

8b.       Pods compressed; flowers yellow, occasionally purplish or pink ............  92. Caragana

7b.       Leaves imparipinnate, rhachis caducous, not hardened.

9a.       Shrubs; flowers in 4–8-flowered raceme; ovary sessile, with stipitate-glandular hairs or villose     93. Calophaca

9b.       Herbs; flowers 1–3; ovary glabrous or villose (including 94. Chesniella in FRPS;

 

 
note! Chesniella = Chesneya, see Pl. Book, if not combined, then see couplet 10) ..........  95. Chesneya

10a.     Stem obvious; stipules membranous, separated from petiole; calyx campanulate; vexillum glabrous .........................................................................................................  94. Chesniella

10b.     Stem abbreviated, stemless-like; stipules herbaceous, lower adnate to petiole; calyx tubular; vexillum densely pubescent below .......................................................  95. Chesneya

6b.       Calyx base symmertric or suboblique; wings usually palmately nerved; pod valves not twisting.

11a.     Leaves 1-foliolate; shrubs; peduncles spine-like ..........................................  100. Alhagi

11b.     Leaves pinnately compound; herbaceous, rarely shrublets or subshrubs; peduncles not spine-like.

12a.     Keel-petals half long as wings; style shorter than ovary or as long as it.

13a.     Style incurved; upper 2 calyx-lobes free; seeds scrobiculate; stipules free and adnate to the base of petiole .................................................................................  98. Gueldenstaedtia

13b.     Style curved in right angle; upper 2 calyx-lobes connate above medium; seeds smooth, marble-maculate; stipules connate below apex and opposite to leaves ..................  99. Tibetia

12b.     Keel-petal subequal or slightly shorter than wings; style longer than ovary.

14a.     Keel-petals obtuse; pod unilocular or with a septum intruding from abaxial suture; leaflets ± symmetrica .......................................................................................  96. Astragalus

14b.     Keel-petals apiculate; pod unilocular or with a septum intruding from adaxial suture; leaflets basally ± unequal-sided or, if narrow, falcately incurved ......................  97. Oxytropis

5b.       Pod-nerves oblique, closely parallel, raised at maturity, confluent towards placental margin; vexillary filament connate with others; keel-petals not auriculate; leaflet-nerves extending to margin (3. subtribe Galeginae) ...................................................................................................  101. Galega

13. Tribe Hedysareae

1a.       Pods not jointed ..........................................................................................  105. Onobrychis

1a.       Pods jointed.

2a.       Pods not prickly ........................................................................................  103. Hedysarum

2b.       Pods prickly on sutures (Note! Stracheya = Hedysarum; see Legumes World, 493. 2005)  104. Stracheya

 

14. Tribe Loteae

Only one genus: Lotus.

15. Tribe Coronilleae

Only one genus: Coronilla.

16. Tribe FABEAE

1a.       Staminal tube with an oblique top; wing-petals adherent to the keel; filaments filiform.

2a.       Style terete; seeds usually more than 2 .................................................................  108. Vicia

2b.       Style dorsally compressed; seeds 2 or solitary ......................................................  110. Lens

1b.       Staminal tube not oblique at the top; wing-petals slightly adherent to or free from the keel.

3a.       Style dilated in the upper part, the inner face longitudinally bearded; filaments filiform or dilated upwards; stipules sagittate or semisagitatte, usually smaller than leaflets ..........................  109. Lathyrus

3b.       Style dilated in the upper part with reflexed margins forming a laterally compressed body; filaments slightly dilated upwards; stipules cordate, larger than leaflets ..........................................  111. Pisum

17. Tribe Cicereae

Only 1 genus: Cicer.

18. Tribe Trifolieae

1a.       Stamens monadelphous; anthers dimorphic; plants usually clothed with glandular and simple hairs       113. Ononis

1b.       Stamens diadelphous; anthers uniform; plants not glandular hairy.

2a.       Petals persisting in fruit; filaments dilated below anthers; pods included in calyx, indehiscent  18. Trifolium

2b.       Petals not persisting in fruit; filaments not dilated; pods not included in calyx, dehiscent or tardily 2-valved.

3a.       Leaves digitately 3-foliolate; flowers solitary or 2–3 in umbels; keel-petals acute  114. Parochetus

3b.       Leaves pinnately 3-foliolate; flowers in racemes, umbels or heads, rarely flowers solitary; keel-petals obtuse.

4a.       Flowers in slender racemes, pod nutlet-like indehiscent or tardily 2-valved
................................................................................................................  115. Melilotus

4b.       Flowers in short racemes, sometimes head-like or solitary; pods straight to coiled.

5a.       Pods straight or curved, linear, long-beaked .........................................  116. Trigonella

5b.       Pods spirally coiled, rarely falcate to nearly straight, often prickly, short beaked  117. Medicago

19. Tribe Crotalarieae

1a.       Perennial herbs; 4 upper calyx-lobes connate in pairs ......................................  121. Lotononis

1b.       Herbs, shrubs or subshrubs; calyx-lobs free or sometimes 2 upper ones connate.

2a.       Pods inflated; leaves simple or 3-foliolate ....................................................  119. Crotalaria

2b.       Pods more or less compressed; leaves 3-foliolate .........................................  120. Priotropis

20. Tribe Eucheresteae

Only one genus: Eucheresta J. Benn.

21. Tribe Thermopsideae

1a.       Stipules partially united and leaf-opposed ......................................................  123. Piptanthus

1b.       Stipules free from each other and not leaf-opposed.

2a.       Evergreen shrubs; bracteoles 2 at middle of pedicel ..........................  124. Ammopiptanthus

2b.       Perennial herbs; bracteoles absent .............................................................  125. Thermopsis

22. Tribe Genisteae

1a.       Herbs; leaves digitately 5-or more-foliolate ........................................................  126. Lupinus

1b.       Shrubs or trees; leaves simple, 3-foliolate or reduced to a scales.

2a.       Small trees; leaves digitately 3-foliolate; pods with thickened sutures ............  127. Laburnum

2b.       Shubs; leaves simple, 3-foliolate, or reduced to a scales.

3a.       Branchlets sharply spiny; leaves reduced to a scales ...........................................  131. Ulex

3b.       Branchlets not spiny; leaves simple, or 3-foliolate, not reduced to a scales.

4a.       Calyx subspathaceous; pods 6–9 cm ........................................................  129. Spartium

4b.       Calyx not spathaceous; pods 1.5–5 cm.

5a.       Leaves digitately 3-foliolate or simple (upper part of plant); pods broad-linear or linear-lanceolate, 2–5 Χ 0.5–1 cm .......................................................................................  128. Cytisus

5b.       Leaves simple; pods linear, 1.5–2.5 Χ 0.3–0.4 cm ....................................  130. Genista